Odyssey of Y Act 4

The Teller at the Gate. The year is 3,800 BCE. A priest-administrator—a "teller"—stands by the oval enclosure at Godin Tepe, assessing and recording goods as they pass through the settlement. A caravan carrying Lapis Lazuli, on its long journey from the mines of Afghanistan to the Uruk civilization of Sumer (modern-day Iraq), unloads its precious cargo for inspection.

As I visualize this scene, I have to wonder: could this teller be my direct paternal ancestor?

We are currently in the Late Chalcolithic (Godin VI), the final chapter before the dawn of the Bronze Age. This is Godin Tepe, a pivotal settlement in the Kangavar Valley of the Central Zagros Mountains. Positioned in what is now western Iran, it served as a high-altitude "dry port" for the ancient world’s most valuable commodities.

Source ©  OpenStreetMaps Modified by myself.

Discussion

I believe that in 3,800 BCE—after many thousands of years—my direct paternal ancestors were likely still situated in the region of the Zagros Mountains. They moved from ibex hunters to gatherers, then to goat herders, and eventually to administrators in a bustling Zagros settlement along the Great Khorasan Road, a vital artery between the Mesopotamian floodplains and Central Asia.

Sites like Godin Tepe have since been rediscovered by archaeologists, proving that these mountains were not inhabited merely by the "savages and barbarians" suggested by early Mesopotamian propaganda. Instead, townships were flourishing within the valleys, especially along these trade routes. These communities were built on a blend of local traditions and the increasing influence of the Sumerian city-states.

While cousins of my direct paternal line radiated eastward across the Iranian plateau toward Gedrosia, Makran, and the Indus Valley—or northward and westward into Anatolia, the Levant, and eventually Europe—my specific line remained. It became what geneticists might call a "ghost" Y-DNA lineage; it never founded a massive, expansive population. The sons carrying L-FGC51040 > FGC51036 never moved in great numbers across Eurasia. Our lineage has always remained rare, perhaps occupying restricted or specialized hereditary roles. Perhaps I truly did have an ancestor who served as a priest or an accountant in a settlement like Godin Tepe.

Odyssey of Y Act 3

The Cradle of the Zagros: 7,500 BCE In a mountain valley in Southwest Asia—modern-day Iran—the Aceramic Neolithic is in full bloom. As I visualize an early goat herder in this landscape, I have to ask: is this man my direct ancestor?

Roughly 9,500 years ago, my paternal lineage (identified by the Y-DNA marker L-FGC51036) likely moved through this transformative agricultural culture. A few millennia later, as the Aceramic period evolved into the Sarab and Guran cultures of the Pottery Neolithic, the next genetic variant appeared: SK1414. Following this mutation, "cousins" of my direct line radiated outward, eventually establishing separate paternal lineages in the Levant, Anatolia, the Indus Valley, and Europe.

My ancestors here were the heirs to accidental selection processes initiated much earlier by the Zarzian culture. Having survived the harsh climatic collapse of the Younger Dryas, these people emerged from their refuges to find their relationship with wild grasses deepening into a state of total interdependence. Over generations, wild flora adapted into domestic strains of einkorn and two-row barley—crops that now relied on human threshing to survive.

Simultaneously, the wild bezoar ibex was being transformed into the world’s first domestic goat. It is highly probable that this region saw the very dawn of goat domestication. These animals were perfectly evolved for the Zagros terrain, turning rugged scrub into high-quality protein for Aceramic farmers. Beyond meat, these goats may have already been exploited for dairy, supplemented by gathered legumes and wild nuts.

However, domestication was neither intentional nor entirely beneficial. While agriculture anchored people to the land and allowed for more children, it also planted the seeds of the global population explosion. In time, humans were conditioned by these new species-to-species relationships. For millennia, early farmers suffered from malnutrition, stunted growth, and zoonotic diseases that jumped from livestock to humans. The status of women often declined; skeletal remains reveal the toll of increased childbirth and the grueling physical labor of milling grain on their knees. Yet, agriculture permits a viral, expansive growth that the old ways could never match.

The Material World of the Aceramic Zagros The inhabitants of the Aceramic Zagros clung to several ancient technologies even as they innovated. They continued to produce delicate flint microliths, though toward the end of the period, they began crafting polished stone axe-heads. They had not yet adopted fired pottery; instead, much like their hunter-gatherer ancestors, they relied on woven baskets and skin bags. Yet, a shift was occurring: they had begun partially firing the clay walls of their storage pits and sculpting small clay figurines. These figures suggest that their belief systems were evolving in tandem with their economy and their changing relationship with the landscape. As visualized in the accompanying images, my ancestors were already fashioning sun-dried clay bricks to build multi-level houses within the Zagros valleys.

Historically, there has been a significant emphasis on the western "leg" of the Fertile Crescent—the Natufians, early Levantines, and Anatolians. However, the importance of this eastern leg in the Zagros has long been underrated. Many population genetics enthusiasts now suggest that Zagros Neolithic Farmers form a distinct genetic cluster that later radiated outward, carrying these foundational agricultural practices with them. I am but one of many descendants of this influential lineage.

Early Fertile Crescent. Home of West Eurasian agriculture.

Source ©  OpenStreetMaps Modified by myself.

Odyssey of Y Act 2

The Zarzian Return: 18,000 BCE I visualize the Zarzian hunter-gatherers of the Zagros and ask: is this my great-grandfather?

The unbearably harsh conditions of the Last Glacial Maximum (LGM) once drove the Baradostian Ice Age hunters out of these mountains and into lower-altitude refuges. However, after 20,000 years ago, the climate began to stabilize, permitting a slow return to the Zagros valleys. Unlike many other groups of the era, my paternal ancestors do not appear to have traveled far; they remained rooted in the Zagros or South Caucasus region for an immense span of time. I would conjecture that they return to the same valleys:

13,000 years ago. Zagros Mountains and valleys (now in Iran) South West Asia

The Zagros and South Caucasus mountains (now in Iran), South West Asia. © OpenStreetMap contributors".

The Zarzian Way: A Prelude to Farming The Zarzian culture was the eastern equivalent of the Epipalaeolithic Natufians in the western Fertile Crescent. While their Baradostian predecessors had sustained themselves primarily by hunting caprines—wild goats and sheep—life after the Last Glacial Maximum shifted toward a broader strategy.

Post-glacial humans enjoyed a far greater diversity of food. Their diet expanded beyond the ibex and mouflon to include onagers, gazelles, crustaceans, fish, birds, and—as visualized above—tortoises. They also gathered a wide array of wild plants, including legumes, nuts, and grass seeds. Evidence of grinding stones suggests they were processing these into porridges or primitive flatbreads.

This period likely represented a critical, early stage in the journey toward agriculture. Accidental selection processes were already in motion. By gathering seeds from plants with firmer rachises (the stem holding the grain), they were unconsciously initiating the shift toward domestication. A similar, unintended process was occurring with their prey. By managing their herds—selecting specific animals to hunt while preserving others—they began the long transition toward animal husbandry. They weren't intentionally trying to "invent" farming; rather, it was nature at work, forging new, symbiotic relationships between humans and the species around them.

Their tools evolved alongside their diet. While the Baradostians excelled at manufacturing burins and regular blades, the Zarzians pushed flint technology further, creating even smaller, geometric microliths designed for complex, composite tools.

Consilience

John Brooker of Long Wittenham, born circa 1720 in Berkshire area? My 6 x great granddaddy. Gemini AI visualised this image based on my own features and on his recorded life.

Why so much hyper-focus on my Y chromosome just recently?  I guess because I have made a few breakthroughs on other lines of the family tree. I had a long conversation with AI about my paternal line and we came to agreement that I had made an error. That had to be put right. After removing those errors, this is where I got back to on my direct paternal line (aka surname line):

  1. My 6x great grandfather was named John Brooker.
  2. My 6x great grandmother was named Mary Gardiner. She was born in 1717 at Hagbourne, Berkshire.
  3. This couple married 1st November 1746 at Oxford College, Oxfordshire. They were both described as residing at Long Wittenham, Berkshire. I have discovered that the reason that they were married at Oxford College was that their parish had been allocated to a vicar living in Oxford College. It was cheaper for them to be married at Oxford, than to call the vicar back to their parish.
  4. Following marriage, John and Mary Brooker proceeded to have at least six children born at Long Wittenham - Mary, Anne, John, Edward, Martha, and a Sarah Brooker. John was a copyhold tenant of St John's College, Oxford, who were one of the two main land holders at Long Wittenham. As a copyholder, John would have cultivated a number of strips in the open-field systems still being used at Long Wittenham. He wasn't a pauper, but neither was he likely to be particularly wealthy.
  5. Before 1746, I have baptism and ancestry for his wife Mary nee Gardiner, but for John, it's the Great Genealogical Dead End.

What were these errors that I deleted for 6 x great grandad John?  I had found numerous John Brookers baptised circa 1720 in the local area. After much deliberation, I settled for the closest village to Long Wittenham, the parish from which his bride had also moved from, I chose a John Brooker baptised at East Hagbourne. But in my heart I knew that there were issues with this choice. For one thing, a lack of correlation in the names of children. Then I became convinced that the John Brooker of Hagbourne, had lived a separate existence from the John Brooker of Long Wittenham. Genealogical crisis!

I immediately recovered from the shock, and started anew with fresh 2026 research. After all, I started out on this quest in 1989, and things have moved on a tad since then. Not only with digitalised, indexed, and online genealogy, but also with genetic genealogy. This is where I shall take this discussion next.

Around 12 years ago, I tested my y-DNA - the DNA that is contained in my Y Chromosome. This can only be passed from biological father to son. Trace it back, and it follows the direct paternal line, all the way back to Y-DNA Adam in Africa. You can use its variants (aka mutations) to tell a story of that one narrow line of descent, way back into prehistory. It does not ALWAYS follow the surname line perfectly, because of illegitimacy as it was previously known, adoption, affairs, and more. People sometimes change name to escape from their past. In the long term, the y-DNA surpasses the origination and adoption of surnames as they occurred during the Medieval.

I've discussed to death the route this genetic evidence tells me that my paternal ancestors must have taken in the past, and will continue to do so:

  1. 28,000 years ago Ice Age hunters of ibex and mouflon in the Zagros mountains of South-West Asia.
  2. 11,000  years ago, hunter-foragers still in the Zagros, or in the Caucasus mountains, who were selectively hunting wild herds, and milling gathered wild seeds in ways that would eventually evolve into the earliest agriculture.
  3. 8,000 years ago my paternal ancestors were aceramic neolithic farmers, herding goats, and sheep. Keeping pigs. Growing emmer and einkorn wheat, barley and more. They may have remained in the area of the Zagros, on its foothills where they founded the ancestry of the Hurrians and Kassites. Or perhaps they had moved immediately southwest, onto the floodplains of early Sumer (Iraq) where they would give birth to a great civilisation?
  4. 4,000 years ago. They would have been in contact with great cities in Mesopotamia, Assyria, Babylonia, the Indus Valley, the Levant.
  5. 3,000 years ago. Perhaps swept westwards to the Levant by the fall of the Mitanni, and the dispersals of the Hurrians.
  6. 600 years ago. Maybe a Genoese or Venetian galley docked at Beirut, Acre, Jaffa or Tripoli and took on my paternal ancestor as crew. My yDNA finally leaves Asia, and heads for Europe.
  7. 600 years ago. A galley docks at Southampton in England, where Genoese and Venetian merchants have a permanent presence, interested in English wool. My last Asian ancestor leaves a son in England.
  8. 400 years ago. Our line has taken the surname Brooker, or is it Chandler? Perhaps a child is born outside of wedlock. He takes his mother's surname. The yDNA is consequently divided among two families in the area of Basingstoke, Kingsclere, Sherfield Upon Loddon, and Newbury across the Hampshire/Berkshire border. Chandlers and Brookers.
  9. 280 years ago. My 6x great grandfather, John Brooker lives in Long Wittenham, Berkshire.

The above is a hypothesis based on those variants on my Y DNA, along with the fact that it survives only in Europe, among two small families who were living only 32 miles apart during the 18th century. Thomas Chandler of Basingstoke, and John Brooker of Long Wittenham. Comparing the DNA, there is a 78% probability that the two families shared the same great grandfather line until circa 1600. Our type of DNA (L-FGC51036) is not found anywhere else in Europe. Our closest Y-DNA cousins are from Iran, and Saudi Arabia. Further back in the variants we have Y-Cousins from Iraq, Lebanon, Pakistan, India, in the Druze ethnicity, in the Parsi ethnicity.

Getting back to the documented record. Our John Brooker. Where did he come from? The DNA does keep pointing south in Berkshire, and to the Hampshire border. A meeting with the Chandlers of Basingstoke. I investigate all of the 'John Brookers' baptised circa 1720 to the south. There can't be many can there? Wrong. I found at least 9 candidates in that part of Berkshire. I have eliminated several from the search, but Berkshire genealogical records are not the best online! I'm usually limited to indexes of transcripts. Transcripts are often wrong. More dead ends.

So I turned to trying to trace back from the Chandlers of Basingstoke hoping that might put me on to the right track for the early Brooker family. I keep hitting more dead ends.

However, here is the thing. Parishes like Chieveley, Newbury, Kingcslere, Sherbourne St John, Sherfield Upon Loddon. I often see entries for baptisms, marriages, and burials for both 'Chandler' and 'Brooker' families. In same villages, registers, sometimes even same pages!

I can smell consilience. Documented trail, and Genetic trail. So close.

Was our Y ancestor a Baloch Lascar?

See page for author [Public domain], via Wikimedia Commons.  "A Portrait of an Indian Gentleman," by A. Smith, 1841.  No.  This portrait was not one of my ancestors.  It is believed to have been painted in England.  However, my Asian Y ancestor must have been here previous to 1700.  This relates to my Y line, inherited down my father's, father's father line and so on.  Descendants for example of Reginald J Brooker, should share this heritage.  My Y-DNA research indicates that I had an Asian ancestor, that most likely moved to Southern England sometime between 1,800 and 500 years ago.  I did find this portrait however, on Wikimedia Commons, whilst searching the subject of who my Y ancestors in Asia were, and why one may have travelled to England.

Let's start a little further back.  My Y-DNA is West Asian in origin.  I share my current terminal Y-DNA SNP (L-SK1414) with a guy that is a Balochi speaker from Makran in SW Pakistan, close to the border with Iran.  I also match fairly well (on STR tests) with a guy who's paternal line hailed from the town of Birjand, South Khorasan, Iran.

Now, although my Australian Y cousin with ancestry in South Khorasan didn't know of any family Balochi link - it's possible.  Balochi, have lived in that region of Eastern Iran.  It may, just may, be a link.  Who are and were the Baloch?

Origins of the Baloch People

See page for author [Public domain], via Wikimedia Commons.  Iranian Baloch khans in Qajar era, c. 1902.  The Baloch people today are spread across Western and Central Asia, mainly found in SW Pakistan, SE Iran, and Afghanistan.  

The pink areas display the main Balochi areas today.  The red outline suggesting likely homelands for my Y-DNA.  Also marked by red spots, are the homes of the two recorded L-SK1414 in that area.  It is estimated that there are 15 million Baloch people across the World today.  The Balochi language is Iranic.  It has been ascribed by linguists as belonging to the North-West Iranic family, close to Kurdish.  Yet the Balochi today, are in the South-East of the Region.  The traditional origin story told by the Baloch people, is that they were Arabic, and originated in Syria.

However, linguists and historians today usually suggest that they were in fact, refugees from Arabic expansion, that migrated mainly east and south east, over several centuries (starting circa 7th Century AD) from an area close to the Caspian Sea in Northern Iran.  This puts Birjand incidentally, on the route of that migration.  It also leads from what I consider to be the homelands of the mother clade - Y hg L1b (L-M317), between the Caspian and Black seas.  Today, the majority of the Baloch are Sunni Muslim (some are Shia).  However, many early migrants from the North West may have ascribed to other religions including Zoroastrianism.  An attack on Persia by the Seljuk Turks from Central Asia during the 11th Century AD, may have accelerated Baloch migration to present day Balochistan. Today, the Baloch of Pakistan, Iran, and Afghanistan are divided into an estimated 130 tribes.

As for Balochistan itself, when Alexander of Macedonia, passed through it, during the 4th Century BC, it was known as the Kingdom of Gedrosia.  Balochistan has long been sandwiched and pulled between the great empires of Persia, and India.  Even today, it is divided between these two political boundaries.  A large region with a sparse population, but a firmly stamped ethnic identity.

According to Akhilesh Pillalamarri "In the 1500s, Balochistan like Afghanistan to its north, became divided into zones of control between the Safavid Persian Empire to its west and the Mughal Empire to its east. This approximately reflects the Iran-Pakistan border today."

Could this friction even have lead my Y ancestor to move?  When did European ships appear on the coastline?

That's the Baloch hypothesis.  Now for the next, the Lascar hypothesis.

The Asian Lascar

By National Maritime Museum from Greenwich, United Kingdom [No restrictions], via Wikimedia Commons. Three lascars on the Viceroy of India.

Our Y ancestor may have moved to Southern England for all sorts of reasons:  merchant, diplomat, adventurer, slave, hostage, prisoner of war, trader, mercenary, servant, religious convert, refugee, etc.  Genetic genealogists tend all too often to cling to known historical events such as battles.  I'd be very wary of that. 

With that in mind... here is one new possibility (as opposed to a probability), that I am presently considering.  

The Lascar.

Lascar derives from al-askar, the Arabic word for a guard or soldier.  When European ships first started to sail the trade routes to India and the Middle East, they often suffered losses of life on the way.  Subsequently, they would recruit new sailors at their ports of call.  Arab traders had scattered seamanship and sailing skills along the coast line around the Persian Gulf, and the practice of Lascars may have already been established before the first Portuguese ships picked them up.  The European practice of taking on Lascars is believed to have started as early as the 16th Century.  It continued through to the 20th Century.  Just about in time, to account for my Y-DNA in Southern England, that turns up during the early 18th Century in two surname families.  It's possible.

Apparently, the Lascars received even poorer food and water than even the late British sailors that they replaced.  Therefore, many jumped ship when they reached England.  Their intentions may not have been immigration, but they couldn't risk the return voyage.  This, it is said, was the very first root of the present day Asian settlement of Britain. It has been speculated that the portrait at the top of the post, may have been of a former Lascar, or ... servant!  Why not though, a traveller that has succeeded?

Why would a 16th or 17th Century European trading ship visit Balochistan?  Did it?  Our ancestor may have already moved either westwards or down to an Indian port.  He may have been a professional sailor!

It's one possibilty.

By National Maritime Museum from Greenwich, United Kingdom [No restrictions], via Wikimedia Commons. Lascars at the Royal Albert Dock in LondonThree lascars on the Viceroy of India. 1936.

Y Haplotype L1b2c

By Hellerick (Own work) [CC BY-SA 4.0], via Wikimedia Commons.  Modified by Paul Brooker.

I've created this distribution map of known Y haplogroup L, L1b2c or L-SK1414. This is my Y-DNA haplotype.  Not a lot of dots there are there?  This is how rare that this clade is.  L1a and L1b most likely (in my opinion) originated during the last Ice Age circa 18,000 years ago, south of the Caucasus, and west of the Caspian Sea in Western Asia.  In other words, in the area of present day Armenia, Azerbaijan, and North-west Iran.  Again, I emphasise, that is just my opinion, looking at present-time evidence.

Y haplogroup L itself may have diverged between L1 and L2, not so much earlier, or so far away from this region.  Again, just my present opinion.

My sub clade of L1b, is so rare, that it is impossible to say.  As can be seen from the map.  However, this is my blog, so I'm going to push out on this one.  My very best guess would be further to the East than it's parent.  I suspect South East of the Caspian Sea, in what is now Eastern Iran.  I could well be wrong.  We have so few tests from nearby Afghanistan for example.  So far, the SNP SK1414 has only been reported twice.  1) in Makran, SW Pakistan, in a Balochi speaking man.  Balochi is an Iranian language, closely related to North-West Iranian languages.  Researchers suggest that the Balochi people of Makran, largely migrated from south west of the Caspian.

The only other guy in the world so far confirmed is little old me, an Englishman.  I trace my surname (direct paternal) line back to the Thames Valley of Oxfordshire / Berkshire 270 years ago.  If my biological line follows that.  A number of STR testers of English descent appear connected to me by STR analysis.  They all descend from Thomas Chandler, who lived around the same time as my earliest recorded ancestor - only 32 miles away at Basingstoke.

From all of the evidence, I conclude that my Y ancestral line moved, probably in one generation, from Western Asia, perhaps from he edge of Persia, to Southern England conservatively between 2,000 and 400 years ago.  Although I would speculate between 1,600 and 600 years ago - during the Medieval or close by.

Y Haplogroup L Resource page

Distribution Haplogroup L Y-DNA

By Crates (Own work) [GFDL (http://www.gnu.org/copyleft/fdl.html) or CC BY-SA 4.0-3.0-2.5-2.0-1.0 (http://creativecommons.org/licenses/by-sa/4.0-3.0-2.5-2.0-1.0)], via Wikimedia Commons.  Unmodified.

Introduction - Y-DNA, Haplogroups, SNPs, Haplotypes

The Y chromosome, and it's Y-DNA, are copied from father to son, down a strictly paternal lineage.  If I were to trace my entire direct ancestry back, I have two parents, four grandparents, eight great grandparents, sixteen great great grandparents.  Yet out of those sixteen great great grandparents (generation 5), who were born only circa 160 years ago, only one carried the Y-DNA that was passed down to me.  My eight great great grandmothers did not not inherit a Y chromosome from their fathers.  Most likely, my other seven great great grandfathers, carried distinctive differently marked Y-DNA.  Yet all sixteen biological great great grandparents have contributed to my overall atDNA (autosomal DNA).  Only one gave me my Y-DNA.  So you can see that Y-DNA represents only one narrow lineage.

Y-DNA, may on the face of it, appear to offer a limited understanding of total biological ancestry.  All sixteen of my great great grandparents were direct ancestors, not just the Y great great grandfather.  However, this lineage offers us evidence that can be genetically tracked, then mapped into relationship.  It could be done to ascertain parental, or non parental events.  It can be used to check the biological validity of relationship to cousins.  As more people investigate and record their haplogroups, haplotypes, STR markers, and SNPs, so we can for example, start to use them to map biological relationship further back.  Y-DNA is particularly useful, not only because of it's markers, but also because it can be plotted to surname studies.  In Western societies, the surname often follows the Y lineage for several generations.

However, Y-DNA (nor the maternal mtDNA) evidence doesn't just stop there.  As more people investigate, submit, and record their data from around the World - and as anthropologists and archaeologists add ancient DNA data from ancient and provenanced human remains to that record, so we can build and plot a world map of the human family, how it relates, how it was distributed globally throughout prehistory.

Both Y and mt DNA carries mutation markers, that define a HaplogroupA haplogroup is a family of shared descent.  These haplogroups are ancient.  The paternal Y-DNA haplogroup that this resource page is dedicated to has been designated as L.

However, mutations do not stop with the formation of a new haplogroup.  They continue through the generations.  As lineages divide between different sons, across many generations, so these mutations in the Y-DNA for example, continue to accumulate down the diverging lineages that once shared common descent.  We are all unique.  The sub clade of L that this page focuses on is L1b.  All male carriers of L1b will carry a SNP (Single Nucleotide Polymorphism) on their Y-DNA that has been designated as M317. This SNP will be downstream of another SNP that has been designated as M22.  Finally, a Y-DNA can be said to have a terminal SNP.  A terminal, refers not to the Haplogroup (in this case L), but can be used to define right down to the last SNP on the Y-DNA, that is shared with others on a record.  If someone for example, carries Y-DNA that is proven (or predicted by comparison) to be Y Haplogroup L, and to carry M317, then their Y terminal could be designated as L-M317, or alternatively, as L1b.  This is also sometimes referred to as a haplotype.  However, a haplotype can also refer to a particular STR.

Y Haplogroup L M20

The above image illustrates a modern day distribution of Y Haplogroup L (M20) as proposed and created by Anthropogenica user Passa.

Y Haplogroup K formed from Y Haplogroup IJK in the Y-DNA of hunter-gatherer fathers and sons, that share a MRCA (most recent common ancestor) during the Upper Palaeolithic, circa 45,400 years ago.  Where did these Y ancestors live at that time?  We think that they lived in Western or Southern Asia.  Iran is a favourite proposal. Earlier Y ancestors had most likely exited Africa 20,000 years earlier, and were well established in Asia.  They had most likely met and confronted another archaic human species, The Neanderthal. This was however, a time of great expansion by humans.  The first anatomically modern humans had recently entered Europe, while other modern humans were arriving in Australia.  The Ice Age was in a flux, but glaciation was advancing.

The most recent common Y ancestor to carry Y Haplogroup LT lived circa 42,600 years ago.  Then a mutation in the Y-DNA lead to the formation of Y Haplogroup L, with a most recent common ancestor 23,200 years ago, close to the time of the Last Glacial Maximum, when ice sheets were reaching their maximum positions.  K, LT, and early L, most likely all originated in Upper Palaeolithic hunter-gatherer populations living during the last Ice Age, in the area of modern day Syria, Iraq, Iran or Pakistan.  It was a time of increased stress on human populations, that were having to adapt to some severe environmental challenges, and may have at times faced isolation into a number of Ice Age Refuges.  Some of these Upper Palaeolithic, Ice Age hunter-gatherer refuges may have been close to the Black Sea, others close to the Caspian Sea, but they were most likely located somewhere between Eastern Anatolia, and Eastern Iran, south of the Caucasus.

L1 / L2 Divergence - the Odd L2's

The oldest divergence within Y Haplogroup L.  L1, as characterised by the SNP M22, diverged from L2, as characterised by the SNP L595.  L2 was only recently discovered, and forced an ISOGG revision of Y Haplogroup L and it's nomenclature that is still causing problems.  In this article, unless stated otherwise, I am using 2017 Nomenclature.  L2 or L-L595 is very rare, but has so far cropped up sporadically across Western Eurasia, including in Azeribaijan, Turkey, Sardinia, England, and Tartaristan.

That is L2 dealt with.  However, most Y Haplogroup L falls into L1. Let us start to look at the main branches of L1.  Remember, L1 is defined by the SNP M22:

Unofficial proposed tree for L1 (L-M22) 2016.  By Gökhan Zuzigo, modified by Paul Brooker.

Proposed Migration Map of L-M22 (L1) by Phylogeographer at https://phylographer.com/mygrations/?

The Big L1 Split - L1a and L1b

As can be seen above, this split occurred around 18,400 years ago, possibly somewhere between what is now Iran and Pakistan.  The L1a branches inherit the SNP M2481, and the L1b branches inherited M317.

First of all, let's look at L1a, because although it is not my sub clade, in terms of modern day population size, it appears to greatly outnumber any other L sub clade.

Pakistan and India - Present Day Home of L1a1 and L1a2

L1a splits again into two sub clades.  The split occurred around 17,400 years ago.  L1a1 (L-M27) and L1a2 (L-M357)

L1a1 (L-M27)

Defined by SNP M27 (on older nomenclature as still used by 23andMe, this was formerly L1*) is mainly found in India, particularly South West India, and in Sri Lanka. This is perhaps the most populous modern day L sub clade, found in 15% of Indian males.  However, it is not restricted to India, and has also been found in 20% of Balochi in Pakistan, and has also been reported in Kirghiz, Pashtun, Tajik, Uzbek, and Turkmen males across Central Asia.

L1a and L1a1 (L-M27) at Birds Eye Cave, Armenia 6161 years before present.

Ancient Y DNA from the Copper Age has emerged from this location in Armenia, and included L1a, and L1a1.  This might suggest, that although very successful today in India and Pakistan, that it has a Western Asian origin.

L1a2 (L-M357)

Has defined by SNP M357 (on older nomenclature as still used by 23andMe, this was formerly L3*).  This sub clade is mainly found in Pakistan, but also Saudi Arabia, Kuwait, The Chechen Republic, Tajikistan, India, and Afghanistan.  It has been found at 15% in Burusho populations, and at 25% in Kalash populations.  It is much more common in Pakistan than in India.

So, the L1a sub clades - spreading down into Southern Asia, and accounting for potentially millions of Y Men there.  Far more than any other branches of Y Haplogroup L.  However, Southern Asia is unlikely to be the origin of L.  That origin is more likely, as stated earlier, to be the place with the most diversity in branches.  That points more towards again towards Western Asia.  It's just that ancient carriers of L, appear to have been particularly successful in Southern Asia, and to have fathered more sons there.

L-M317 or L1b of Western Asia

We now move onto the branches of particular interest to myself, because I carry a Y Haplotype that belongs here.  L1b is defined by the SNP M317, that formed circa 18,400 years ago, most likely in the area of modern day Iran, or elsewhere in Western Asia.

Phylogenetic tree of L1b by Anthrogenica user Caspian (with permission):

Click on above hyperlink for full sized image

L1b is mainly distributed across Western Asia, from modern day Turkey, across to Pakistan.  However, as we will see, it also spreads in low densities across parts of Europe.  it is very much, the "Western L".

The Next split - L1b1 or L-M349.  The Levant, and Europe!

Around 14,000 years ago, another split occurred in the L1b (M317) branch. A new SNP, M349, defined L1b1.  Today, L1b1, or L-M349, is found in Western Asia, in Lebanon, Syria, Turkey, Armenia, etc.  However, it is also found scattered in low densities through parts of Europe.  It crops up in South Europe, often close the the Mediterranean Sea, including particularly in parts of Italy.  It also forms a light cluster in Central Europe.

A working map of Y haplogroup L sub clades by Edward Chernoff.  This map is incomplete, but is published here with permission of Edward Chernoff.  Copyrights applied.

Branching away from a common Y ancestor with L1b1 (M349), is another 14,000 year old line defined by SNP SK1412, L1b2.

L1b2 (L-SK1412) splits - Pontic Greeks, and the others...

13,000 years ago,, during a cold stage towards the end of the last Ice Age, the L1b2 (SK1412) Y branch divides again.  Very recent research suggests that it split into three lines: L-SK1415 (L1b2a), L-PH8 (L1b2b), and L-SK1414 (L1b2c).

L1b2a (L-SK1415), has as far as I know, only been detected in a Makrani Balochi survey in SW Pakistan.

L1b2b (PH8), is found in Turkey, Greece, Armenia, Chechen Republic, Iraq, etc.  It is associated particularly with the Pontic Greek ethnicity from Eastern Anatolia, and around the Black Sea.  A further division within PH8 has been detected at around 3,000 years ago.

Finally ... mine:

L1b2c (L-SK1414, FGC51074), has so far been SNP detected only in Makrani Balochi, in SW Pakistan, Gujarat, India, Turkey, Cyprus, Saudi Arabia, Lebanon (Druze), and in England. STR predictions for L-SK1414 have also been found in Goa, Syria, Iraq, Kuwait, UAE, Tartaristan, France, Italy, Iran, and Azores.  In addition to SK1414, I have with the assistance of Gareth Henson, a FT-DNA Big Y test, accompanied by further analysis of their raw data, by Yfull, and FullGenomes, ascertained 117 novel SNPs looking for first time matches.  As can be imagined, I'm very keen that further L Y-Men should test.

Distribution may be connected to the dispersal of the recently identified group known as the Iranian Neolithic Farmers after 10,000 BCE:

https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5113750/#:~:text=We%20sequenced%20Early%20Neolithic%20genomes,significantly%20to%20the%20ancestry%20of

In addition there would appear to be a potential link between this group, and the inhabitants of the Harappan or Indus Valley civilization. Below is my proposed distribution of L SK1414 / FGC51074:

Those tentative European Y haplogroup L links

We have seen above, that again, and again, Y haplogroup L (M20), and several of it's sub clades appear to have Western Asian origins, despite success of some of those sub clades today in India and Pakistan.  Y haplogroup L has not been linked to the Yamna hypothesis, that has taken credit for the origin of many haplogroups that are successful today in Europe.  Y-DNA L was located to the southern side of the Caucasus, between present day Turkey and Pakistan.  However, two particular Y-DNA L sub clades do make mysterious appearances across Europe.

1) L-L595 (L2) has only recently been discovered, so far, exclusively across Europe, in very low numbers.

2) L-M349 (L1b1), downstream of M317, also spreads across South Europe, and clusters at the Rhine-Danube.  I have on 23andMe forums, seen a number of testers that unfortunately have not tested their Y elsewhere, claim Ashkenazi paternal ancestry, but this is far from common to all European L-M349 samples. Although rarely forming much more than 1% of all Y along the Mediterranean coast of Southern Europe, this percentage does occasionally rise higher, for example, in parts of Italy.

When did L2 or even L1b1 enter Europe?  L2 has only so far been found in Europe.  There are some suggestions that some European L could be survivors from the Eurasian Neolithic.  However, ancient DNA has not yet been found to support this hypothesis. 

Prime resources

L Yfull Tree

https://www.yfull.com/tree/L/

Wikimedia entry for Y Haplogroup L-M20

https://en.wikipedia.org/wiki/Haplogroup_L-M20

FTDNA L The Y Haplogroup L Project

https://www.familytreedna.com/public/Y-Haplogroup-L/

Marco Cagetti's Y Haplogroup L

http://www.cagetti.com/Genetics/L-haplogroup.html

Anthropogenica Y Haplogroup L Forum Board

http://www.anthrogenica.com/forumdisplay.php?37-L

ISOGG 2009 Y Haplogroup L (Useful for understanding 23andMe Y haplogroup result of L2*)

http://isogg.org/tree/2009/ISOGG_HapgrpL09.html

ISOGG 2017 Y Haplogroup L

http://isogg.org/tree/ISOGG_HapgrpL.html

Other resources

Europedia Y-DNA Haplogroup L

http://www.eupedia.com/europe/origins_haplogroups_europe.shtml#L

23andMe users should note that the company in 2016, still used a very outdated ISOGG nomenclature system.  My 23andMe reported haplotype was L2*.  However, using ISOGG 2016, this is now L1b (L-M317).  NOT to be confused with modern day L2 (L-L595).

Facebook Y Haplogroup L Group

https://www.facebook.com/groups/773887796013634/

L-M317 STR Alpine cluster article

https://figshare.com/articles/L_M317_STR_marker_likelihood_tree_focuing_Alpine_cluster/105684

Familypedia Wiki for Haplogroup L

http://familypedia.wikia.com/wiki/Haplogroup_L_(Y-DNA)

For personal note  as of 2018-08-28.

My Y Haplogroup L Designation:

L +M20 +M22 +M317 +SK1412 +SK1414 (or FGC51074) +FGC51041 (or Y31947) +FGC51036

L-SK1414 = L1b2c

SK1414/FGC51074 age estimate current 9,300 years bp.

L-FGC51041 is a verified terminal

FGC51041/Y31947 age estimate current 6,000 years ago but only 2 samples on Ytree.

L-FGC51036 on terminal on FT-DNA

L-Y31947 is terminal on yFull

115 novel SNPs