In 2015, the Peopling of the British Isles (POBI) research group, published this paper:

The Fine-scale genetic structure of the British Population  Leslie, Winney etal POBI 2015

It proposed that the Early Medieval migration events commonly known as Anglo-Saxon (a better term to include the 9th Century surge could be Anglo-Danish), has been exaggerated. They concluded that the modern English had only 10% to 40% descent from these Continental immigrants, with the remainder majority reflecting earlier Iron Age / Romano British ancestry.

An independent 2016 investigation by Schiffels, Haak etal looked at ancientDNA from cemeteries in Cambridgeshire. The results supported POBI's conclusion, proposing:

'East English population derives 38% of its ancestry from Anglo-Saxon migrations'

• POBI 2015 suggested 10% - 40% Anglo-Danish
• The small scale Schiffels, Haak etal report of 2016 suggested 38% 'Continental Northern European'  (Anglo-Danish) in the Cambridgeshire region.

The 2022 study based on hundreds of ancient remains increased the percentage of new arrivals. They conclude that it is higher further east, closer to the North Sea, but declines as an average in Western England.  At its peak in Eastern England, they projected that Anglo-Danish accounted for 76% of the genome:

'the individuals who we analysed from eastern England derived up to 76% of their ancestry from the continental North Sea zone, albeit with substantial regional variation and heterogeneity within sites.'

This is a higher estimate than that proposed by the previous two studies.

Image of an LBA socketed axehead from Portable Antiquities Scheme

The Forgotten Contribution to the South British Genome.

 A research team looking at ancientDNA led by David Reich had already detected a 97% population replacement across Britain at the close of the Neolithic period, circa 2,300 BCE. They proposed that the Earlier, European Neolithic Farmers of Britain, were replaced by a new people, associated with the Bell Beaker artefact culture. These new people had previously been admixed between European Neolithic Farmers living on the Continent, and recently migrants from the Pontic & Caspian Steppes of Eurasia. 

Northern Europe has more of this migrant Yamnaya / Steppe ancestry which arrived during the 3rd Millennium BCE, while Southern Europe (peaking in Sardinia, then Iberia) has more residual ancestry from the earlier, European Neolithic Farmers. Yet the South British (English) have rather more Neolithic ancestry than other Northern neighbours. This raised questions concerning where had this DNA come from.

Reich's team speculated on this result, and investigated the remains of ancient DNA further. This was later reported on:

Large-scale migration into Britain during the Middle to Late Bronze Age

https://eprints.whiterose.ac.uk/183814/2/Manuscript.pdf

In the above paper, the research team suggest a secondary migration event, that followed the Bell Beaker population replacement. They date it to the end of the Middle Bronze Age / start of the Late Bronze Age circa 900 BCE, although proposed that it had slowly been arriving for some time, before a surge of new arrivals to Britain. They do not pinpoint where these immigrants come from, but by their heavier Neolithic ancestry, it is proposed that they had moved up from further south, most probably from France. How many? The study proposes a 50% DNA replacement in Southern Britain, across England & Wales. I think that is probably comparable to the most recent, highest estimates for the much later Anglo-Danish immigration event.

The tabloids of course, reacted:

From this it has further been proposed, that it may have been this forgotten immigration which brought the p-celtic and / or q-celtic languages to Britain. If you subscribe to identifying Iron Age Britons as Insular Celts then this could represent the arrival in Britain. 

Personally I feel that what we 21st Century CE people believe to be Insular Celtic, reflects a much longer, older exchange of people and ideas across Britain, Ireland, and the Western Seaboard of Europe during the Bronze and Iron Ages. The Romans did much later, claim more recent Belgae migrations into Britain, some tribes even shared names with tribes in France and Belgium. These could represent a continuation of migration, possibly of elites. Prehistoric Britain was very much in contact with the nearby Continent, and a part of Europe.

Conclusion

Original Nature publication:

https://www.nature.com/articles/s41586-021-04287-4

On BBC:

https://www.bbc.co.uk/news/science-environment-59741723

Earlier Posts:

The Beaker phenomenon and genetic transformation of Northwest Norfolk. A layman's take 2017

https://paulbrooker.posthaven.com/the-beaker-phenomenon-and-genetic-transformation-of-northwest-europe-2017-a-laymans-take

Celebrating my Steppe and Beaker ancestors

https://paulbrooker.posthaven.com/celebrating-my-steppe-and-beaker-ancestors

Own Photo.

Comparing results from actual recorded ancestry, to that predicted by Ancestry.com, 23andme, My Heritage, Living DNA, FT-DNA and more.

Recorded Ancestry

Results

Lola, Mesolithic girl on National Geographic

The above is a link to a reconstruction by Tom Björklund of a Mesolithic girl who lived in what is now Denmark. It is a very unique and creative reconstruction, because nothing physical of this girl nicknamed Lola survives into our archaeological record! She is only known by her DNA (and that of recently eaten food) that she left on a lump of birch tree pitch that she had chewed as a gum some 5,700 years ago. But I really like this reconstruction. I think that she makes a beautiful wild child. Straight out of the lines of a novel that I'm trying to write.

Analysis of her DNA strongly suggests brown toned skin, if not dark brown. Her hair dark brown. The DNA supports that her eyes were light coloured. Perhaps blue, blue-green or hazel? I'll return to Lola, but first these features correspond to those suggested by the genomes of other Mesolithic remains scattered around Europe.

Cheddar Man who lived in South West Britain around 10,000 years ago is the best known. The revelation several years ago that the DNA sequenced from his genome, suggests both dark skin and blue eyes caused quite a commotion. A lot of people didn't like it, and accused the geneticists of woke.

We always knew that the earliest modern humans were likely to have plenty of melanin. Subsisting on a hunter-gatherer diet that was rich in dietary Vitamin D meant that there was little adaptive pressure for them to lose this dark skin in a hurry. Just as some hunters in the far north and far south have retained dark skins into recent times. The emphasis to reduce melanin may not have arrived until following major shifts to a poor, agricultural diet in northern zones. The DNA associated with very light skin of modern Western Europeans may not have arrived until quite recently (prehistorically speaking).

We also had Villabruna Man in Northern Italy. His presence and DNA is less known to the general public. But even before the controversy of the Cheddar Man reconstruction, we knew from Villabruna and other remains that he lacked certain genetic indicators of light skin and:

Additional evidence of an early link between west and east comes from the HERC2 locus, where a derived allele that is the primary driver of light eye color in Europeans appears nearly simultaneously in specimens from Italy and the Caucasus ~14,000-13,000 years ago.

https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4943878/

But one recent study proposes that there were at least two distinct clusters of hunter-gatherers around Europe at the close of the last Ice Age:

On the basis of the genetic variation of present-day Europeans, this could imply phenotypic differences between post-14 ka hunter-gatherer populations across Europe, with individuals in the Oberkassel cluster possibly exhibiting darker skin and lighter eyes, and individuals in the Sidelkino cluster possibly lighter skin and darker eye colour.

Imagination

I recently had my Ancestry.com / Ancestry.co.uk results updated in the beta test - for myself and for my family.  The new results make AncestryDNA my most accurate DNA test so far.  Here are the screenshots for the latest results for my family kits:

My results before the Update:

Following the latest 2018 update:

My sibling's new results:

My Mother's:

I've updated my spreadsheet comparing different results for myself, from different vendors in order to reflect how well that the new Ancestry test is now working for myself and my family, compared to 23andme etal;

A few more screenshots:

During the Black Friday sales last December, I bought two Ancestry kits. I actually mean't to order one, but made a bit of a mess of it. Still, I thought, they were as cheap as I've seen a DNA kit, so I let the order process.

Some people might exclaim - but you've tested your DNA to death!  These kits were not directly intend for myself though.  I intended them for the art of Genetic Genealogy.  To help me verify my paper tree, biologically.

Although I enjoy the kick that I get from matching segments of DNA in strangers, to shared ancestors of the past on all lines, I'm particularly interested in one line, from one great grandfather.  You see, I had a very naughty great grandmother.  I have uncovered evidence of two bigamous marriages by her, as well as other relationships.  A second cousin of mine, through her, doesn't appear to have the amount of shared DNA that I would expect for a full second cousin.  It looks worse than even the old family rumours.

As I do have an extensive family tree down to that birth certificated great grandfather, even though I know full well that biological family isn't always as good family as non-biological, which the paper trail honours, I'd still like to know.  With Genetic Genealogy, I hope to verify - or otherwise, his biological relationship.

So... I used one kit to test one of my siblings, and the other to test my mother.  I've tested my mother before on 23andme.  Mistake.  I've learned a lot about DNA testing over the past few years or so.  Ancestry.com might seem like a heavy marketing, greedy big DNA company, with some slightly dishonest sales ploys (find out if your ancestor was a Viking!), and pressure to subscribe to more services in order to get the full benefit of the test - BUT ... it 1) has an awesome family tree building website for subscribers, that link to DNA tests, 2) has the largest customer database, and 3) through it's genealogy services, as well as marketing, has the most UK testers in it's database.

Okay, it's a little dumbed down.  The messaging system sucks (so I always send my email address), It doesn't provide a chromosome browser.  It doesn't provide segment locations on chromosomes.  But - for my uses - using DNA matches to verify a family tree pedigree, it serves extremely well.  I have had almost ten times more matches on AncestryDNA, than from 23andme, FT-DNA, and GEDmatch combined.  And many have online trees!

I've received my siblings results.  Wow.  I suspected it.  That the sibling has inherited some quite different DNA from the parents mean't that although we share some DNA matches, there are many that we don't!  Up to now, I've just used a spreadsheet to keep results of verified matches.  I could see that I now need something more powerful.  Something that I could search on - and filter different lineages.  When my mother's results arrived, I'll be able to divide all of my matches into maternal, or paternal sides.  On top of that, I have a 1C1R (first cousin once removed) on my father's side, that I can sometimes use to indicate some ancestry on his side.  I can look at all of my matches and their shared matches, and triangulate, where abouts they fit into my family tree.  I built a personal database for my DNA matches.

So I'm pretty pleased that I invested in those two kits during the sales.  It's kept me busy.

I used Open Office Base to build the database:

Okay it's basic and not pretty, but I can extend on it.  I've imputed our closest 187 DNA matches, nearly all from Ancestry, plus a few verified from FT-DNA and GEDMATCH.  It's a family match - I've included forms for imputing my mother's and sibling's matching segments - not just my own.  Any genuine matches that my sibling has - are also my cousins.  Just that I don't have personally share DNA segments with them.  I've also included a yes/no check box for that 1C1R.

I've used it to query an up-to-date list of "our" shared DNA matches that share a correlating common ancestor or two on their trees with ours.  My biological "verifiers".

Using the open source GRAMPS app, I produced a fresh family pedigree fan chart.  I then used open source GIMP to colour in the ancestors that I have verified with shared DNA segments.  The darker the tone, the more matches:

It's generally looking pretty verified isn't it.  My birth certificate grandparents were all very clearly, my biological grandparents.  The great grandparents, and the majority of great great grandparents are also looking pretty verified.  But what about that great grandfather?  The birth certificate version was my surname great grandparent, and biological version was my Y-DNA great grandparent.  Were they the same?

Well I still do not have evidence that I'd regard as overwhelming.  But I am gathering evidence that he may have been the same guy.  I have two DNA matches that strike directly through him.  Unfortunately, both were distant ancestry, with only a small shared segment each (around 7 cM).  That small, they could either belong to an undocumented relationship elsewhere, or even be identical but not by descent.  But it's evidence that I'm building, and it's more reassuring than if he'd had no DNA matches strike through his lineage to us.  The other supportive evidence was that my biological paternal line great grandfather carried an incredibly rare haplogroup: Y haplogroup L-SK1414 (L1b2c).  The only other L-SK1414 so far found in the British Isles, traced his paternal surname line back to Basingstoke, around 1740.  My documented surname line traces back in 1740 to Long Wittenham, Berkshire.  Only about 32 miles away from the Basingstoke L-SK1414 by road.  Could be a coincidence, but it supports that the Y-DNA could still correspond with the surname line back in 1740, and that my great grandaddy, was my DNA great grandaddy.

Such is the power of genetic genealogy.  Roll on the results of my mother.  That will reduce the number of matches that are likely to be on my paternal side.

But I'm afraid I don't buy it.

The above image illustrates of some of my ancestral locations, as according to documented genealogy.  As can be seen.  I have quite a lot of East Anglian ancestry.  What might also be observed, is the location of East Anglia, and of South East England, in relation to Belgium, the Netherlands, and North East France.

I'm an East Anglian.  Much of my family tree is East Anglian.  Before documented genealogy picks up my family trail, who were the East Anglians, what were their origins?  The traditional answer would be that they were the descendants of the Angles.  An early 5th Century AD tribe, that relocated from Angeln, now in Schleswig Holstein, on the North Germany, South Denmark border.  Them, and maybe a few Saxons, Jutes, Suevvi, etc.  All pretty much from what is now North Germany and Denmark.  See the map below:

Archaeology sort of backs this up ... but also offers some slight alternatives.  At first, British Archaeology supported the Historians - that there was a near genocidal event during the 5th and 6th centuries, where the Anglo Saxon tribes arrived and displaced the Romano-Britons that had until then, lived in South East Britannia.  There certainly is plenty, even, overwhelming evidence, of Anglo-Saxon culture if not settlement in East Anglia at this time.  Below are the locations of a few of the many Anglo-Saxon cemeteries found in East Anglia - and their closest artifact correlations on the Continent - in Northern Germany:

It's all adding up.  However, then, a new trend appears in British Archaeology that plays down the Anglo-Saxon invasion hypothesis. From the 1980's onward, some British archaeologists started to argue that they saw patterns of land use continuity between the Romano-British and Pagan Saxon periods.  They argued there was no archaeology of genocide.  No battle sites.  No mass graves.  Instead they proposed that only limited numbers of Anglo Saxons arrived - and that their culture was largely adopted by the Romano-Britons that already lived here.  Some even suggested that no Anglo Saxons came here - it was merely a cultural import.

Then Genetics stepped in.  Most notably with POBI (Peopling of the British Isles) 2015, but also with a number of other studies, often comparing the DNA from excavated remains to modern populations.  They proposed a new middle house consensus.  No there was no genocide.  The modern English have more old British ancestry than Anglo-Saxon.  However, there was a significant Anglo Saxon immigration event.  But they mixed, intermarried.  Anglo-Saxon culture was adopted, but Anglo Saxons had not displaced the Britons.  They had married them.  The modern English it seems have around 10% to 40% Anglo Saxon ancestry, and 60% to 90% British.  Sort of watered down Celts.

The assumption that we have made, is that rural populations on the front-line immigration - such as East Anglians, were the most watered down, with highest percentages of Anglo-Saxon ancestry.  Why not.  The archaeology would support that.  The East Anglian landscape is littered with Anglo-Saxon, and Anglo-Danish place-names.  It is the most Anglo-Saxon landscape, and later, was firmly within the Dane-Law.

My Ancestry

It's all over this blog.  However, in summary, I am an East Anglian.  Born at Norwich, with all four grandparents of Norfolk birth.  I have been researching my family tree using genealogy, for thirty years, on and off.  I have accumulated the recorded names of 490 direct ancestors.  Around 80% or more of this ancestry lived here in East Anglia.  At Generation 6 (3rd great grandparent), my ancestry was 97% South East English, and 3% Swiss.  This is demonstrated in this fan chart of my recorded ancestry:

If that's not East Anglian enough - look at the right hand side of that fan chart, my mother's side.  My mother has 225 of her direct ancestors recorded, and everyone lived firmly in East Anglia.

I have documents, likenesses in family photos, and family stories to back my narrative ancestry; but I am also gradually building biological evidence through the use of DNA matches to other testers, that share a common ancestry with me, that correlates well with the shared DNA:

That the vast majority of our ancestry is very rural, and poor agricultural working class, would suggest that we have had ancestry here in East Anglia for a very, very, long time.  I have traced some lines back to early parish records in the 16th Century.  I would expect that many of our ancestors belonged to peasant families in Medieval Norfolk and Suffolk.  These, I'd expect were the descendants of Anglo Saxons, Romano-Britons, and Danes.

Here comes the paradox.

Population Genetics and the DNA

Documentary evidence confirms that I'm an East Anglian, and English.  But when I tested with the commercial DNA-for-ancestry vendors, such as 23andme or FT-DNA, my results, although seeing me as pretty firmly, a North West European, doesn't really see me as particularly British.  23andme suggested only 32% British.  They instead suggest that my ancestry is rather Continental.  High levels of "West European" or "French & German".  I at first assumed that this was ancient or early medieval ancestry, my Anglo-Saxon and Anglo-Danish East Anglian roots showing through.

But is that the case?  Analysis using some of the latest calculators challenge that.  Here are the PCA locations of myself and my mother, on one such calculator.  This one (By David Wesolowski) includes many modern references grouped on language family, but also includes some references from actual ancient DNA in Northern Europe, including some extracted from Anglo Saxon remains in Cambridgeshire, close to East Anglia:

You see the red squares represent the Anglo Saxons in Cambridgeshire?  Mine and even my very rural East Anglian mother's place well to the right of them, closer to modern day Irish speakers, overlapping with modern day French speakers.  More Celtic it seems than Anglo Saxon.

Here's another PCA showing our positions (red myself, orange, my mother), this one by Lukasz, but based on the Eurogenes K36 calculator:

It puts me closer to Flemish then Walloon, followed by SE English.  Finally a third PCA, from Eurogenes K15: 

We consistently position between SE England and North France, Belgium, and the Netherlands; rather than between SE England and Northern Germany and Denmark.  It appears that we relate closer to Normans, Belgians, Walloons, Flemish, and Dutch - than we do to modern day North Germans or Danes.  Here is our K36 Oracle maps by Lukasz:

My mother has a slight more pull from Denmark and Schleswig Holstein.  Perhaps this is from early medieval Angle/Danish settler in East Anglia?  However, we both pull strongest outside of England, from the Low Countries.  Flemish, Dutch, and Walloon come up as closest Continental matches.  I'm not surprised.  I've noticed for some time, that the big vendors appear to give some testers of Normandy, Hauts-de-France, Belgium, and the Netherlands ancestry, very similar results to my own.

If our results were at all representative of East Anglians of local ancestry, then the modern East Anglian is perhaps as much, or more of a Belgian than he or she is a Dane or Angle.  So I've added a secondary circle to the map that I used above, to illustrate what are popularly beleved to be the origins of the East Anglians.  The new blue circle, represents not what history or archaeology suggests, but what my family's DNA currently suggests:

Note that I have included South East Britain in there - because clearly, there was no displacement.  The Romano-Britons were among our ancestors.

The Language Connection.

It has long been noted, that the very closest dialect or language to English, is West Frisian.  Old Frisian and Old English (or Anglo Saxon) were close.  Linguists group them together as "Anglo-Frisian".  Why is this?  If we all descend from Angles and Danes?

When did our "Belgian" ancestors arrive in Britain?

By Belgian, I'm referring to ancestors that we share not only with Belgians of local ancestry, but also the North French, and the Dutch.  Answer.  I don't know.  All that I am pointing out, is that the DNA of my East Anglian family appears to be more like that of people that today live in that part of the Continent, than in Denmark, Norway, or even North Germany.  However, here are a few ideas:

1. The Bell Beaker.  During the Late Neolithic.  We believe that the British Bell Beaker people largely crossed over from the Lower Rhine Valley.
2. The Belgae.  I'm not quite so sure about this one, but let's just go with it.  Roman historians recorded a late Iron Age migration from the Belgium area, into South East Britain.  They described them as using a Celtic language and culture, but being closer related to Germanic tribes to the east.
3. An unrecorded migration from Northern France to Southern Britain during Late Prehistory.  This one was suggested by POBI 2015, that claimed to detect a relationship between the Southern British and Northern France, that they claimed was most likely Late Prehistoric.
4. Roman Britain.  Britannia was often administered along with Gaul.
5. Saxo Frisia.  Saxons didn't only Southern Britain, they also settled the Low Countries, where they took the name of the earlier tribe there - the Frisians.  Did many Anglo-Saxon settlers actually crossed over from Frisia?
6. Norman.  1066 and all that.  A whole new elite arrived, often bringing artisans and supporters with them.
7. Angevin and Medieval French.  For a time, large regions of France were ruled along with England.  French artisans, merchants, monks, priests, etc.
8. The Elizabethan Strangers.  Protestant refugees were invited from the Low Countries, both Dutch and Walloons.  They particularly settled towns in South East England such as Norwich, and Colchester during the 16th and 17th centuries.
9. The Huguenots.  French Protestant refugees that arrived during the 17th and 18th centuries.
10. Background migration.  My favourite.  In addition to all of the aforementioned proposed migration events, the slow, gradual contact between South East England and the Low Countries / Northern France, that has always been there.  The drip-drip in the background.  That the North sea and Dover Strait separating the two areas is so narrow.  Merchants, refugees, masons, artisans, weavers - the Dutchmen and Frenchmen recorded as Aliens in many post medieval surveys.  The fishermen from Haut-de-France that frequently beached on the Norfolk coast.  The dutch herring fishermen that traded herring at Yarmouth market.