Gravettian - into the Big Freeze

Image Source. The Venus of Brassempouy.

The Gravettian overlaps the late Aurignacian, dating from circa 33,000 years ago, and surviving until 20,000 years before present. This included the coldest peak of the last Ice Age, the Last Glacial Maximum around 24,000 years ago until 18,000 years ago. These really were Ice Age Europeans. Travel as far north as Britain must have taken place during warmer intervals. Otherwise the Gravettian is found in a band across Western Eurasia, which stretches from Portugal and the Basque region, through France, Germany, Czech republic, as far east as Georgia and South Russia.

Image Source. Female face. Ivory carving, Dolní Věstonice.

Gravettian yDNA haplogroups so far discovered are CT, I, IJK, BT, one C1a2 and one F.
Gravettian mtDNA haplogroups overwhelmingly U (mainly U2 and U5), with one M.

The Late Gravettian developed into a culture in France and Iberia, that we call the Solutrean. Meanwhile in the south east of Europe, from the Italian peninsula across the the Western Steppe, it diverged into the Epigravettian culture, which overlaps for much of its range with the Mammoth Steppe.

We identify a genetic ancestry profile in individuals associated with Upper Palaeolithic Gravettian assemblages from western Europe that is distinct from contemporaneous groups related to this archaeological culture in central and southern Europe4, but resembles that of preceding individuals associated with the Aurignacian culture. This ancestry profile survived during the Last Glacial Maximum (25,000 to 19,000 years ago) in human populations from southwestern Europe associated with the Solutrean culture, and with the following Magdalenian culture that re-expanded northeastward after the Last Glacial Maximum. Conversely, we reveal a genetic turnover in southern Europe suggesting a local replacement of human groups around the time of the Last Glacial Maximum, accompanied by a north-to-south dispersal of populations associated with the Epigravettian culture. 
A previously unknown population replacement event that was archaeologically invisible, occurring during the peak of the Ice Age.  This DNA was then passed on via the Solutrean people to the following Magdalenian people. 

Image Source. Triple Burial at the Upper Palaeolithic Site of Dolní Věstonice.

These people buried their dead more frequently in graves than did the Aurignicians.  The grave of three male teenagers at Dolní Věstonice was particularly enigmatic. The person on the left, has a hand placed over the stomach of the middle character, where red ochre had been applied. The central person had a curved spine, diagnosed as caused by chondrodysplasia calcificans punctata (CCP).  The body on the right had been placed facing down. Red ochre had also been applied to the heads of all three.

A Mirror into the Past and Present. On the Dolni Vestonice Triple Burial. Academia.edu 2020. Olga Viviana Nauthiz Szynkaruk
For their last journey, the individuals have been richly equipped with headdresses made of the teeth of the arctic fox (Vulpes lagopus), wolf (Canis lupus), and unidentified large predators. In the mouth of DV15, a mammoth ivory disc was placed.

Image Source. Gravettian points.

Prey appear less restrictive than for the earlier period which focused on reindeer. The Gravettians hunted deer, mammoth, hares, lions, bear, foxes. Rather than use split antlers as points, they used flint blades known as Gravettian points. There is evidence that the Gravettians had access to bows and to the spear thrower. Furthermore, they may have had the first domesticated animal, the dog.

Image Source (Flickr).  Altamira Cave Paintings. By Carlos Calamar

British Gravettian

Image Source (modified).  Find-spots in Britain of Gravettian tanged points. Northern Britain was covered by ice sheets, and the North Sea was dry, connecting Great Britain to the Continent.

Image Source. The Venus of Willendorf.

Conclusion

The Gravettians entered Europe in the long, slow run up to the Last Glacial Maximum, and endured much of it. Although they inhabited a vast region of Western Eurasia, their culture remained constant, albeit with some divisions between west and east. The Pavlovian Culture developed in the east as mammoth-hunters.  The Gravettians often buried their dead with much ceremony, wearing seashell beads, ivory, and animal teeth. They were successful and adaptive Ice Age hunters, preying on deer, reindeer, mammoth, fox, hare, hyena, wolf, seal, and foraged for shellfish. They lived in circular, semi subterranean shelters, but were mobile. Skilled artists with cave paintings and carved ivory. They tipped projectiles with flint blade points, and may have used bows, boomerangs and atlatls. Nets were woven, and lamps of stone used. They may have developed an early relationship with dogs.

During the Last Glacial Maximum, although their artefact culture continued, there was a population replacement event by people closer related to their Aurignician predecessors.  These later Gravettians fostered the Solutrean Culture in France and Iberia, and the Epigravettian culture of the east and south east.

Aurignacian - Reindeer hunters and first modern artefact culture of Europe

Image of the Lion Man.  The Oldest Portable Art: the Aurignacian Ivory Figurines from the Swabian Jura (Southwest Germany).  Harald Floss

The Aurignacians were not the first Europeans.  With the previous post on the Apidima 1 skull fragment from Greece, dated to circa 210,000 years ago, I established that humans with modern Homo sapiens features may well have been wandering in and out of parts of Europe for a very long time. Neither was Apidima 1 the first European. Earlier humans, including Neanderthals had been around Europe for a very long time. Before them, earlier hominins, such as Homo heidelgergensis; and Homo antecessor who left artefacts and footprints on a Norfolk beach, some 800,000 to 900,000 years ago. Recently, stone tools found in Ukraine during the 1970s have been dated to 1.4 millions years of age, and may be associated with an Homo erectus type hominin.

Not the first Europeans, but here in this post I am going to investigate the earliest modern human artefact culture that we currently know to have established itself in Europe, and even in Britain. I'm going to discuss the Aurignacians.

Image.  Animation and Graphic Narration in the Aurignacian. Marc Azéma

Thought to have spread into Europe from SW Asia in the Levant, where the culture is also found, it has been proposed that an earlier origin could be the Zagros Mountains of Western Iran, where similar tools have been recorded.

Aurignacian yDNA haplogroups so far discovered are C1a, C1b, and K2a
Aurignacian mtDNA haplogroups include N, R, and U.

Of these, only the mtDNA hapologroup U is still common in modern Europe.

The genetic history of Ice Age Europe. Nature 2016. Fu, Posth, Hajdijak etal.  (Full download here)

Concluded that a 37,000 year old Aurignacian genome has some continuity into the modern European population, and was more akin to modern Europeans, than was a contemporary sample from China. The division between Western Eurasians, and Eastern Eurasians dates back to include the Aurigacians in the West. A contribution to modern European DNA has been identified albeit a small percentage. The genomes sequenced indicated that they were likely dark-skinned and brown eyed, but with reservation.

Here we present genome-wide data from three individuals dated to between 45,930 and 42,580 years ago from Bacho Kiro Cave, Bulgaria1,2. They are the earliest Late Pleistocene modern humans known to have been recovered in Europe so far, and were found in association with an Initial Upper Palaeolithic artefact assemblage. Unlike two previously studied individuals of similar ages from Romania7 and Siberia8 who did not contribute detectably to later populations, these individuals are more closely related to present-day and ancient populations in East Asia and the Americas than to later west Eurasian populations. This indicates that they belonged to a modern human migration into Europe that was not previously known from the genetic record, and provides evidence that there was at least some continuity between the earliest modern humans in Europe and later people in Eurasia. Moreover, we find that all three individuals had Neanderthal ancestors a few generations back in their family history, confirming that the first European modern humans mixed with Neanderthals and suggesting that such mixing could have been common. 
Recent arrivals into Europe, with connections to present day West Eurasian populations, and they had some recent Neanderthal ancestry mixed into modern.

According to Wikipedia:
The Proto-Aurignacian and the Early Aurignacian stages are dated between about 43,000 and 37,000 years ago. The Aurignacian proper lasted from about 37,000 to 33,000 years ago. A Late Aurignacian phase transitional with the Gravettian dates to about 33,000 to 26,000 years ago.
That is actually a very long time period. One research project proposed a Europe-wide population of only 1,500 at a time. Extended over a long period of many thousands of years.

The early Aurignacian dispersal of modern humans into westernmost Eurasia. PNAS 2020. Haws, Benedetti, Talamo and Zinsious.

Looks at the entry into Iberia, and revises the date.

Image. Antler points and a perforated baton from the Early Aurignacian.  Origin and Development of Aurignacian Osseous Technology in Western Europe: a Review of Current Knowledge  Élise Tartar

Interview with Dr James Dilley on the use of these antler points. He suggests that a lack of good wood with which to construct spear shafts may have led to them employing more breakable split antler points to preserve the valuable shafts. The points could have also improved the bleeding out of quarry. Perforated batons are also common on their sites. Use is unknown.


Flint bladelets as with some later cultures, are often a feature of Aurignacian sites. Typical hunter's lithics utilising flint with great economy.

Continental sites produce volumes of reindeer bone. It seems to have been their target prey.

Their landscapes were increasingly cold, open, and treeless.

Culturally they left high quality cave paintings in South France, and carved ivory pieces in Germany. The first Venus figure. The Lion Man. They often painted and sculptured lions, which may have been important to their belief system.

Image of the Venus of Hohle Fels.  The Oldest Portable Art: the Aurignacian Ivory Figurines from the Swabian Jura (Southwest Germany).  Harald Floss

British Aurignacian

Image. Recorded Aurignacian sites in Britain.  Coastline would have been far further out than in the above image. The North Sea was dry, and Britain connected to the continent.

The Timing of Aurignacian occupation of the British Peninsula.  Edinburgh Research Explorer 2012. Dinnis.R

A study of flint burins to type in comparison with sites in Belgium and France. Occupation of the British peninsular would have been impossible for long periods. The conclusion:

British Aurignacian burins busqués are technologically indistinguishable from those found in Belgium and at Abri Pataud in southern France c. 32 000 14C BP, or c. 37 000 cal BP. Therefore, the Aurignacian can be considered to have appeared in Britain at this same time. The proposed c. 32 000 14C BP appearance of burins busqués accords with the few radiocarbon dates from other sites which directly date Aurignacian occupation of Britain. Morphologically similar lozangic-type osseous points are also present at Abri Pataud and in Britain at this time. This period apparently coincides with or closely follows the most significant warm phase during the lifetime of the Aurignacian: Greenland Interstadial 8. An environmental response to this climatic amelioration is therefore a plausible reason for the extension of Aurignacian ranges northwards at this time.

and:
In spite of an overall paucity of material, the presence of two bladelet production techniques suggests that there were at least two Aurignacian occupations of Britain, or that occupation was sufficiently prolonged to encompass the replacement of one by the other. The precise timing of what is interpreted as the more recent of the two techniques – the Paviland burin method – is currently unknown.
More than one occupation during warmer periods around 32,000 years ago, or / and 37,000 years ago. These coincide with warmer interstadials. Find-sites include Goughs Cave, Kents Cavern, and Goats Hole, Paviland. Britain's classic Aurignacian skeletal remains are those of The Red Lady of Paviland. A male who had died in Britain circa 31,000 years ago.

Image of the Aurignacian flute made from vulture bone.  The Oldest Portable Art: the Aurignacian Ivory Figurines from the Swabian Jura (Southwest Germany).  Harald Floss

Video. Hear the bone flute being played.

Conclusion

Ice Age reindeer hunters on the European tundra with a talent for the arts. They hunted with split antler tipped throwing spears. They had music, and made flutes, using the long leg bones of vultures. They were talented artists, leaving ivory and bone sculptures, and their famous cave paintings.  They were few, and moved around far, following herds and shifts in the bitter weather. Their landscape was open and cold, treeless. Fauna would have included reindeer, tarpan / horses, steppe bison, woolly rhino, mammoth, cave lions, and hyenas. The lion may have been ritually important to their belief systems. They most probably encountered another type of human in Europe - the Neanderthals. Four percent of their own DNA, with long segments, originated among the Neanderthals.  The artefact culture survived for thousands of years, until the approach of the Last Glacial Maximum some 25,000 years ago. They persist in a few percent of modern West Eurasian DNA.

In addition to reindeer hunting, some sites are associated with ancient coastlines, and pierced seashells have been found as personal ornamentation. No evidence of fishing, but they may have foraged for shellfish. Ornamentation also includes the teeth of carnivores such as lions and foxes. Red ochre was applied on some remains.


This investigation has really helped me to imagine them. The Europeans who lived here before Last Glacial Maximum. I was really surprised just how many resources there are available online. I've barely touched on this subject. I have only touched on their cave art

I don't want to violate copyright by sharing Tom Björklund's fantastic art work here, but here is a link to his take on the Aurignacian people. I think that creativity blended with archaeology really helps:

Notes for Mesolithic Western Europeans. The last free people of Europe.

Lola, Mesolithic girl on National Geographic

The above is a link to a reconstruction by Tom Björklund of a Mesolithic girl who lived in what is now Denmark. It is a very unique and creative reconstruction, because nothing physical of this girl nicknamed Lola survives into our archaeological record! She is only known by her DNA (and that of recently eaten food) that she left on a lump of birch tree pitch that she had chewed as a gum some 5,700 years ago. But I really like this reconstruction. I think that she makes a beautiful wild child. Straight out of the lines of a novel that I'm trying to write.

Analysis of her DNA strongly suggests brown toned skin, if not dark brown. Her hair dark brown. The DNA supports that her eyes were light coloured. Perhaps blue, blue-green or hazel? I'll return to Lola, but first these features correspond to those suggested by the genomes of other Mesolithic remains scattered around Europe.

Cheddar Man

Cheddar Man who lived in South West Britain around 10,000 years ago is the best known. The revelation several years ago that the DNA sequenced from his genome, suggests both dark skin and blue eyes caused quite a commotion. A lot of people didn't like it, and accused the geneticists of woke.

We always knew that the earliest modern humans were likely to have plenty of melanin. Subsisting on a hunter-gatherer diet that was rich in dietary Vitamin D meant that there was little adaptive pressure for them to lose this dark skin in a hurry. Just as some hunters in the far north and far south have retained dark skins into recent times. The emphasis to reduce melanin may not have arrived until following major shifts to a poor, agricultural diet in northern zones. The DNA associated with very light skin of modern Western Europeans may not have arrived until quite recently (prehistorically speaking).

We also had Villabruna Man in Northern Italy. His presence and DNA is less known to the general public. But even before the controversy of the Cheddar Man reconstruction, we knew from Villabruna and other remains that he lacked certain genetic indicators of light skin and:

Additional evidence of an early link between west and east comes from the HERC2 locus, where a derived allele that is the primary driver of light eye color in Europeans appears nearly simultaneously in specimens from Italy and the Caucasus ~14,000-13,000 years ago.

https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4943878/

But one recent study proposes that there were at least two distinct clusters of hunter-gatherers around Europe at the close of the last Ice Age:

On the basis of the genetic variation of present-day Europeans, this could imply phenotypic differences between post-14 ka hunter-gatherer populations across Europe, with individuals in the Oberkassel cluster possibly exhibiting darker skin and lighter eyes, and individuals in the Sidelkino cluster possibly lighter skin and darker eye colour.

https://www.nature.com/articles/s41586-023-05726-0

 I think that may be close to the truth, that there was most likely a variation in skin tones and eye colours across Europe during late prehistory.

Then we met 'Elba the Shepherdess' from Galicia, Spain. She dates to around 9,300 years ago. Analysis of Elba's ancient DNA suggests that she was dark-skinned and haired, and brown-eyed. Her remains had spectacularly been excavated along with three aurochs (wild cattle). Her relationship to these aurochs has raised extremely controversial questions about possible domestication. I do like the aurochs. I might make them the a subject of a later post.


Imagination


British Archaeology has long been going through a struggle where it tries to lose its roots within the Arts, and relocate itself within the (social) sciences. Clinging vigorously to data, they fear telling any stories (which is what hi-story does). In my own creative writing, I have my British hunter-gatherers at the close of the Mesolithic as shameless animists, who see themselves very much as a part of Nature. Mine are trapped into nations or tribes, divided into semi-nomadic bands who wander a larger region that I call a wilderness. In the background (but no longer in the novel) from their own golden age, they look back through folklore to an earlier time when they were more free, to wander further, one nation, following herds of steppe and forest bison.

In their own contemporary wilds 6000 years ago, I have them hunting red deer, roe deer, wild pig, aurochs (wild cattle), red squirrel, martens, seal, porpoise, whales, beaver, fox, waterfowl (ducks/geese), bustards, cranes, wood pigeon, woodcock, fishing/trapping eel, salmon, trout, chub, pike. Foraging for hazelnut, acorns (which they need to process to reduce tannin), cat-tails, wild garlic, pig-nuts, harvesting wild grass seeds, tubers, roots, tree sap, flower buds, lichens, sea lettuce, samphire, berries (blackberry, raspberry, elderberry, hawthorn), black bee honeycomb (yes they make berry mead and alcoholic birch sap), greens (including some springtime tree leaves), tree inner bark (in desperation or to chew), fungi (bollettes, blewitts, ceps, chanterelle, puffballs, chicken-of-the-woods and many more), mussels, sea molluscs /shellfish, crustaceans. I did also include beechnuts, but then found that the beech may have only just arrived with the Neolithic. Ugh! 

One thing that this creative writing has helped me understand about the British Late Mesolithic, is that the forests at that time lacked a wealth of biodiversity. Britain had parted from the European Continent too soon following the glaciers - Ireland more so. Few tree species had made it here. The temperate wild-woods (I don't think all of those in the south-east could have been classed as rain-forests) could be very mean with any calories between early winter and mid spring. I emphasise that in the story, the need to gather in late summer and through the autumn, and to process and store what they could. Acorns and hazel nuts could be roasted and even ground into a flour to make breads.  Wild grass seeds may have been added. The game would have been fatter at that time of year, and the salmon would run. Oh, I dare to suggest that my hunter-foragers are also gardeners of their wilds, encouraging hazel and birch to spread and survive.

Late winter may have been comparatively miserable, where the bands would laze close to hearths. Conserving valuable calories of their energy.

I enjoy imagining the wild-woods of SE Britain 4000 BCE. Perhaps not all temperate rain-forest, but neither anything like a modern day woods. Deadwood, flooding, saplings, rot, deep ferns and mosses. They would have been difficult to pass through by foot. Waterway would have been preferred. Trees with mosses and octopus boughs. Lime (linden) trees, elm, wych elm, oak, birch, alder, willow, ash, pine.  Wolves, lynx, brown bear. Eagles, black woodpeckers, and goshawks. But I don't imagine it all as wild-wood. I have opted in my imagined Britain 4000 BCE, for lots of small glades, and larger open plains that in the story, I call prairies. Here herds of aurochs join those of red and roe deer, to keep this scrub and grassland opened up. Bustards parade by orchids, while the cuckoo calls.

But all in my imagination. Not fact.



Returning to Lola in Denmark. The gum also revealed that she was lactose intolerant, and this has also been found in other genetic remains of this period. The DNA of hazel and mallard duck was also on the gum, and it is thought that hazelnut and duck may have been her recent meals. I think it is incredible archaeology that a lump of birch pitch could produce such a result. The 5,700 ybp was based on radio carbon dating of the lump of Mesolithic chewing gum.

When I was a voluntary archaeologist and field walker, I would treasure any microliths, microlith waste cores (I had a few), and a tranchet axe head (below) that I found from the Mesolithic. It was always my favourite period of prehistory. When people would have related to other species as a part of the Natural World to which they belonged. Innate animism that I strongly relate to as an autistic biophilliac. Or did they? Next post I will look at Gobekli Tepe and other sites of this same period across Anatolia. Have we simplified the savage?


So for all of their imperfections and inaccuracies, here are the faces of the Mesolithic. The last truly free wild people of European Nature. The last savages of Europe. Before we started to screw it all up.  Lola is my favourite.

My Basal-rich K7 Results

David Wesolowski of the Eurogenes Blog, has created a new ancient admixture calculator, the Basal-rich K7.

In his blog, he states: "The Basal-rich K7 is the best ancient ancestry test that I've been able to come up with. It correlates strongly with latest research reported in scientific literature. And, in fact, in some instances it probably trumps latest scientific literature.

For instance, Broushaki et al. 2016 characterized Early Neolithic farmers from the Zagros Mountains, Iran, as 62% Basal Eurasian and 38% Ancient North Eurasian-related (Figure S52). This, considering formal statistics like the D-stat below, with AfontovaGora3 (AG3) as the ANE proxy, is unlikely to be correct, despite the fact that AG3 is a relatively low quality sample.".

Villabruna-related

The Villabruna cluster represents the DNA found in 13 individuals in Europe from after 14,000 years ago.  They were Late Ice Age hunter-gatherers.  They appear to have links with the Near East.  The current thought is that they replaced earlier groups of hunter-gatherers in Europe.  The DNA of people in the Middle East and Europe pulled together at this time, and they may represent an expansion from the South-East.  Much of the Aegean Sea would have been dry, with low sea levels (glaciation), so the migration may have been easy.  It is believed that they had dark skin, and blue eyes.  They were possibly, the last hunter-gatherers of Europe and the Middle East.  They may have contributed to our DNA both through or either, later Asian or European admixtures.

David gives the English average as 56.7%.  My result is 57.1%

Basal-rich

The Basal Eurasians are a hypothetical "ghost" population derived from DNA studies.  It is suggested that they splintered from other modern humans 45,000 years ago, presumably outside of Africa, somewhere around the Middle East.  They significantly contributed DNA to the Early Neolithic Farmers of the Fertile Crescent and Anatolia, and consequently, on to all of us modern West Eurasians.  

 David gives the English average as 26.5%.  My result is 28.8%

Ancient North Eurasian

Another Ice Age hunter-gatherer "Ghost" population, but this one has been associated with human remains and an Upper Palaeolithic culture (Mal'ta-Buret') at Lake Baikal, Siberia.  We know that it significantly contributes to modern West Eurasians, through earlier admixture on the Eurasian Steppes.  Copper Age pastoralists then carried it westwards into Europe with their later expansion.

David gives the English average as 16.6%.  My result is 14.0%

Others

David gives the English averages as SE Asian 0.15, Oceanian 0.07, East Eurasian 0.00% and Sub Saharan 0.00

My results are SE Asian 0.00, Oceanian 0.01, and Sub Saharan 0.05

Comparison with other testers

A remarkable similarity has been observed between many of my East Anglian atDNA results and a Norman tester.  On K7, we are almost identical.  Indeed, we are often closer to each other in results, than I am to other British, and he is to other French.

I'm increasingly recognising that although my East Anglian heritage should in theory bring me closer to North German and Scandinavian results, in practice, compared with other Britons, I am pulled more to the south - to France, and even to Southern European.  Hence, I tend to receive lower ANE than many British, Irish, or Scandinavian, and more Early Neolithic Farmer in ancient admixture tests, than would be expected.

Other than Norman admixture, I struggle to explain this with either known paper recorded ancestry (252 direct ancestors from East Anglia and SE England - 100% English), or with known regional history.